Nathan Fillion (born March 27, 1971) is a Canadian-American actor, best known for the leading roles of Captain Malcolm "Mal" Reynolds on Firefly and its film continuation Serenity, Richard Castle on Castle and John Nolan on The Rookie. Fillion has acted in traditionally distributed films like Slither and Trucker, Internet-distributed films like Dr. Horrible's Sing-Along Blog, television soap operas, sitcoms and theater. His voice is also featured in video games, such as the Bungie titles Halo 3, Halo 3: ODST, Halo: Reach, Destiny and Destiny 2, along with the 343 Industries video game Halo 5: Guardians.Fillion first gained recognition for his work on One Life to Live in the contract role of Joey Buchanan, for which he was nominated for a Daytime Emmy Award for Outstanding Younger Actor in a Drama Series as well as for his supporting role as Johnny Donnelly in the sitcom Two Guys and a Girl. Nathan Fillion was born in Edmonton, Alberta, Canada, the younger of two sons of Robert "Bob" Fillion and June "Cookie" Fillion (née Early), both retired English teachers. Both sides of his father's family were part of the Quebec diaspora in Fall River, Massachusetts, and his mother had a Norwegian maternal grandfather and a Finnish maternal grandmother. Fillion was raised in Edmonton's Mill Woods neighborhood and completed his secondary and post-secondary education in Edmonton, attending Holy Trinity Catholic High School, Concordia University College of Alberta, and the University of Alberta, where he was a member of the Kappa Alpha Society. He has been a U.S. citizen since 1997.

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