Ashley Miller was born on March 16, 1971 in Windber, Pennsylvania, Miller's family relocated, and he grew up in Gainesville, Virginia. His first job out of college was as a middle school English teacher. He then went on to work for the United States Navy as an independent defense contractor. Miller eventually moved to California, starting his career as a writer on the television series Andromeda after meeting one of the producers of the series. He is best known for his work on the television series Andromeda, Terminator: The Sarah Connor Chronicles and Fringe. He also worked on the films Thor and X-Men: First Class. He often collaborates with fellow screenwriter Zack Stentz.

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