Helen Quilley (Gwyneth Paltrow) gets fired from her public relations job. As she leaves the office building, she drops an earring in the lift and a man picks it up for her. She rushes for her train on the London Underground and misses it. The plot then splits into two storylines, presented as parallel universes, in which different events ensue, consequent upon whether or not she catches a particular train.In the timeline in which she boards the train, Helen sits alongside James (John Hannah) (the man in the lift) on the Underground and they strike up a conversation. She gets home in time to catch her boyfriend, Gerry (John Lynch), in bed with his American ex-girlfriend, Lydia (Jeanne Tripplehorn). Helen dumps him and moves in with her friend Anna (Zara Turner), and, at Anna's suggestion, she changes her hairstyle to make a fresh start. James continues to serendipitously pop into Helen's life, cheering her up and encouraging her to start her own public relations firm. She and James fall in love despite her reservations about beginning another relationship so soon after her ugly breakup with Gerry...
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